A wildlife photographer has captured a series of spectacular shots of birds of prey at a north-east nature reserve.
A group of five to seven owls have taken up residence at Scottish Natural Heritage’s Forvie reserve, near Collieston, since last October.
Ron Macdonald, who is a former SNH manager, has been monitoring the birds for four months and managed to get some spectacular images of them swooping on prey.
He said: “Short-eared owls are a dream to photograph. Because they fly in daylight, often slowly and perch on posts, they make fantastic portraits.
“They can be tricky to photograph when flying against the background of heath and dunes, but with some basic field craft and photo skills, you will soon get the hang of it.
“They also become more comfortable with people over the winter, becoming familiar with the visitors hoping to capture their good looks and their interesting behaviour.”